Recent Species

Genus: Paralulworthia
Class: Sordariomycetes Subclass: Lulworthiomycetidae
Order: Lulworthiales
Family: Lulworthiaceae

Genus: Paralulworthia
Class: Sordariomycetes Subclass: Lulworthiomycetidae
Order: Lulworthiales
Family: Lulworthiaceae

Genus: Annabella
Class: Leotiomycetes
Order: Helotiales
Family: Cordieritidaceae

Genus: Roussoella
Class: Dothideomycetes
Order: Pleosporales
Family: Thyridariaceae, Subclass Pleosporomycetidae

Genus: Pontoporeia
Class: Dothideomycetes
Order: Pleosporales
Family: Halotthiaceae, Subclass Pleosporomycetidae

Genus: Neocamarosporium
Class: Dothideomycetes
Order: Pleosporales
Family: Neocamarosporiaceae Wanas., Wijayaw., Crous & K.D. Hyde, Studies in Mycology 87: 245 (2017), Subclass Pleosporomycetidae

Genus: Morosphaeria
Class: Dothideomycetes
Order: Pleosporales
Family: Morosphaeriaceae, Subclass Pleosporomycetidae

Genus: Kirschsteiniothelia
Class: Dothideomycetes
Order: Pleosporales
Family: Kirschsteiniotheliaceae, Subclass Pleosporomycetidae

Genus: Halobyssothecium
Class: Dothideomycetes
Order: Pleosporales
Family: Lentitheciaceae, Subclass Pleosporomycetidae

Genus: Falciformispora
Class: Dothideomycetes
Order: Pleosporales
Family: Pleosporaceae, Subclass Pleosporomycetidae

Recent Publications on Marine Fungi

New publications

Poli A, Bovio E, Ranieri L, et al (2020) Fungal diversity in the Neptune Forest: Comparison of the mycobiota of Posidonia oceanica, Flabellia petiolata, and Padina pavonica. Front Microbiol.

Gonçalves MFM, Santos L, Silva BMV, Abreu AC, Vicente TFL, Esteves AC, Alves, A. (2019) Biodiversity of Penicillium species from marine environments in Portugal and description of Penicillium lusitanum sp. nov., a novel species isolated from sea water. Int J Syst Evol Microbiol 39, 3014–3021.

Gonçalves MFM, Esteves AC, Alves A. (2020) Revealing the hidden diversity of marine fungi in Portugal with the description of two novel species, Neoascochyta fuci sp. nov. and Paraconiothyrium salinum sp. nov. Int. J. Syst. Evol. Microbiol.: 10.1099/ijsem.0.004410, 6 (2020)

Gonçalves MFM, Silva BMV, Esteves AC, Alves A (2019) Verrucoconiothyrium ambiguum sp. nov., a novel species isolated from sea water, and affiliation of the genus Verrucoconiothyrium to the family Didymellaceae. Int J Syst Evol Microbiol, 69(12):3769-3776 (2019)

Fotedar R, Sandoval-Denis M, Kolecka A, Zeyara A, Al Malki A, Al Shammari H, Al Marri M, Kaul R, Boekhout T (2019) Toxicocladosporium aquimarinum sp. nov. and Toxicocladosporium qatarense sp. nov., isolated from marine waters of the Arabian Gulf surrounding Qatar. Int J Syst Evol Microbiol, 69(10):2992-3000 (2019)

Marchese, P., Garzoli, L., Gnavi, G., O’Connell, E., Bouraoui, A., Mehiri, M. et al (2020) Diversity and bioactivity of fungi associated with the marine sea cucumber Holothuria poli: disclosing the strains potential for biomedical applications. J Appl Microbiol 129: 612-625.

Abdel-Wahab MA, Jones EBG, Bahkali AH (2020) Marine fungi recorded from Avicennia marina (Forsk.) Vierh. and their secondary product potential. Nova Hedwigia 111 (3-4): 357–390

Brandon T. Hassett & Rolf Gradinger (2018) New Species of Saprobic Labyrinthulea (=Labyrinthulomycota) and the Erection of a gen. nov. to Resolve Molecular Polyphyly within the Aplanochytrids. Journal of Eukaryotic Microbiology 2018, 65, 475–483

Brandon T. Hassett (2020) A Widely Distributed Thraustochytrid Parasite of Diatoms Isolated from the Arctic Represents a gen. and sp. nov. Journal of Eukaryotic Microbiology 2020, 0, 1–11 1

Fryar SC, Hyde KD, Catcheside DEA (2020) A survey of marine fungi on wood in South Australia. Bot Mar 63: .

Fryar S, Hyde KD, Catcheside DEA (2020) A checklist of marine fungi from Australia. Mycotaxon 135:465–465.

Nagano Y, Miura T, Tsubouchi T, Lima AO, Kawato M, Fujiwara Y, Fujikura K (2020) Cryptic fungal diversity revealed in deep-sea sediments associated with whale-fall chemosynthetic ecosystems. Mycology 11:263–278 .

Su CJ, Hsieh SY, Chiang MWL, Pang KL (2020) Salinity, pH and temperature growth ranges of Halophytophthora isolates suggest their physiological adaptations to mangrove environments. Mycology 11:256–262 .

Abd El-Rahman TMA, Tharwat NA, Abo El-Souad SMS, El-Beih AA, El-Diwany AI (2020) Biological activities and variation of symbiotic fungi isolated from Coral reefs collected from Red Sea in Egypt. Mycology 11:243–255 .

Hagestad OC, Andersen JH, Altermark B, Hansen E, Rämä T (2019) Cultivable marine fungi from the Arctic Archipelago of Svalbard and their antibacterial activity. Mycology 11:230–242 .

Luo Y, Wei X, Yang S, Gao YH, Luo ZH (2020) Fungal diversity in deep-sea sediments from the Magellan seamounts as revealed by a metabarcoding approach targeting the ITS2 regions. Mycology 11:214–229 .

Tibell S, Tibell L, Pang KL, Calabon M, Jones EBG (2020) Marine fungi of the Baltic Sea. Mycology 11:195–213 .

Gleason FH, Allerstorfer M, Lilje O (2020) Newly emerging diseases of marine turtles, especially sea turtle egg fusariosis (SEFT), caused by species in the Fusarium solani complex (FSSC). Mycology 11:184–194.

Devadatha B, Calabon MS, Abeywickrama PD, Hyde KD, Jones EBG (2019) Molecular data reveals a new holomorphic marine fungus, Halobyssothecium estuariae, and the asexual morph of Keissleriella phragmiticola. Mycology 11:167–183 .

Ettinger CL, Eisen JA (2020) Fungi, bacteria and oomycota opportunistically isolated from the seagrass, Zostera marina. PLoS ONE 15(7): e0236135.

Ettinger CL, Eisen JA (2019) Characterization of the mycobiome of the seagrass, Zostera marina, reveals putative associations with marine Chytrids. Front Microbiol 10: 2476.

Vohník, M; Borovec, O; Kolařiková, Z; Sudová, R; Réblová, M. 2019. Extensive sampling and high-throughput sequencing reveal Posidoniomyces atricolor gen. et sp. nov. (Aigialaceae, Pleosporales) as the dominant root mycobiont of the dominant Mediterranean seagrass Posidonia oceanica. MycoKeys. 55:59-86

Marine Microbial Diversity as a Source of Bioactive Natural Products
Didier Stien (editor)
MDPI • Basel • Beijing • Wuhan • Barcelona • Belgrade
Interesting chapter on marine mycology techniques.
Didier Stien
Marine Microbial Diversity as a Source of Bioactive Natural Products, Reprinted from: Mar. Drugs 2020, 18, 215, doi:10.3390/md18040215

David P. Overy, Teppo Rämä, Rylee Oosterhuis, Allison K. Walker and Ka-Lai Pang
The Neglected Marine Fungi, Sensu stricto, and Their Isolation for Natural Products’ Discovery, Reprinted from: Mar. Drugs 2019, 17, 42, doi:10.3390/md17010042 .

Young Mok Heo, Kyeongwon Kim, Seung Mok Ryu, Sun Lul Kwon, Min Young Park, Ji Eun, Kang, Joo-Hyun Hong, Young Woon Lim, Changmu Kim, Beom Seok Kim, Dongho Lee and Jae-Jin Kim
Diversity and Ecology of Marine Algicolous Arthrinium Species as a Source of Bioactive Natural Products, Reprinted from: Mar. Drugs 2018, 16, 508, doi:10.3390/md16120508

Anna Poli, Elena Bovio, Lucrezia Ranieri, Giovanna Cristina Varese and Valeria Prigion (2020). News from the Sea: A New Genus and Seven New Species in the Pleosporalean Families Roussoellaceae and Thyridariaceae. Diversity 2020, 12, 144 (1-18) ; doi:10.3390/d12040144

Dayarathne M. C., Wanasinghe D. N., Devadatha B., Abeywickrama P., Jones E. B. G., Chomnunti P., Sarma V. V., Hyde K. D., Lumyong S. & McKenzie E. H. C. 2020.— Modern taxonomic approaches to identifying diatrypaceous fungi from marine habitats, with a novel genus Halocryptovalsa Dayarathne & K.D.Hyde, gen. nov. Crypto gamie, Mycologie 41 (3): 21-67. http://

Jones EBG, Devadatha B, Abdel-Wahab MA, Dayarathne MC, Zhang SN, Hyde KD, Liu JK, Bahkali AH, Sarma V V., Tibell S, Tibell L, Wang MM, Liu F, Cai L (2020) Phylogeny of new marine Dothideomycetes and Sordariomycetes from mangroves and deep-sea sediments. Bot Mar 63: .

Jones EBG, Pang KL, Abdel-Wahab MA, Scholz B, Hyde KD, Boekhout T, Ebel R, Rateb ME, Henderson L, Sakayaroj J, Suetrong S, Dayarathne MC, Kumar V, Raghukumar S, Sridhar KR, Bahkali AHA, Gleason FH, Norphanphoun C (2019) An online resource for marine fungi. Fungal Divers 96:347–433 .

Pasqualetti M, Giavannini V, Baragjini P, Gorrasi S, Fenice M (2020) Diversity and ecology of culturable fungi associated with Posidonia oceanica leaves and their epiphytic algae Dictyota dichotoma and Sphaerococcus coronopifoilius. Fungal Ecol. 44: 100906

Dayarathne MC, Jones EBG, Maharachchikumbura SSN, Devadatha B, Sarma VV, Khongphinitbunjong K, Chomnunti P,  Hyde KD (2020) Morpho-molecular characterization of microfungi associated with marine based habitats. Mycosphere 11(1): 1–188.

Chuaseeharonnachai C, Suetrong S, Nuankaew S, Somrithipo S, Hongsanan S, Srikitikulchai P, Jones EBG, Boonyuen N. (2020). Synnematotriadelphia gen. nov. (S. stilboidea comb. nov. and S.  synnematofera comb. nov.) and Triadelphia hexaformispora sp. nov. in the family Triadelphiaceae. Mycological Progress, 19:127-137.

Dayarathne MC, Wanasinghe DN, Devadatha B, Abeywickrama P, Jones EBG, Chomnunti P, Samra VV, Hyde DK, Lumyong S, McKenzie EHC (2020) Modern taxonomic approaches to identifying diatrypaceous fungi from marine habitats with a novel genus Halocryptovalsa Dayarathne & K. D. Hyde, gen. nov. Cryptogamie, Mycologie 41 (1): 1-47.

Sheng-Nan Zhang, Mohamed A. Abdel-Wahab, Kevin D. Hyde, E.B. Gareth Jones, Ratachadawan Cheewangkoon, JIAN-KUI LIU (2019). Additions to the genus Savoryella (Savoryellaceae), with asexual morph Savoryella nypae comb. nov. and S. sarushimana sp. nov. Phytotaxa 408(3): 195-207.

Sanja Tibell, Leif Tibell, Ka-Lai Pang and E.B. Gareth Jones (2019). A conspectus of the filamentous marine fungi of Sweden. Botanica Marina. (pp 1-14)

Ariyawansa HR, Jones EBG (2019) Additions to Taiwan fungal flora 2: Ophiosphaerella taiwanica sp. nov. Phytotaxa413 (1): 039–048.

 Brandon T. Hassett, Tobias R. Vonnahme, Xue-fang Peng, E.B. Gareth Jones, Céline Heuzé (2019). Global diversity and geography of the planktonic marine fungi. Botanica Marina (pp pp 1-20)

Dayarathene MC, Maharachchikumbur SSN, Jones EBG, Al-Sad AM, Hyde KD, Chomunti P (2019) Sexual morph of Phaeoacremonium aureum from Rhizophora mucronata collected in southern Thailand. Phytotaxa 387 (1): 021–039

Ekanayaka AH, Jones EBG, Hyde KD, Q. Zhao Q (2019). A stable phylogeny for Dactylosporaceae. Cryptogamie, Bryologie 40 (3): 23-44.

Zhang SN, Hyde KD, Jones EBG, Jeewon J, Cheewangkoon R, Li JK (2019) Striatiguttulaceae, a new pleosporalean family to accommodate Longicorpus and Striatiguttula gen. nov. from palms. MycoKeys 49: 99-129

Devadathaa B, Calabon MS, Abeywickramaab PD, Hyde KD and Jones EBG (2019). Molecular data reveals a new holomorphic marine fungus, Halobyssothecium estuariae, and the asexual morph of Keissleriella phragmiticola. Mycology: An International Journal on Fungal Biology10.1080/21501203.2019.1700025.



Abdel-Wahab MA, Jones EBG, Abdel-Aziz FA, Bahkali AH (2019) Nia lenicarpa sp. nov. (Niaceae, Agaricales) from Red Sea mangroves in Saudi Arabia with comments on Nia vibrissa. Phytotaxa 406(3): 157-168.

Calado M, Carvalho L, Barata M & Pang KL (2019) Potential roles of marine fungi in the decomposition process of standing stems and leaves of Spartina maritima. Mycologia 111(3): 359-370.

Letcher PM & Powell KJ (2019) A taxonomic summary of Aphelidiaceae. IMA  Fungus 1(4): 1-11 .

Nahano Y, et al. (2019) Deep-sea endemic fungi? The discovery of Alisea longicolla from artificially immersed wood in deep sea off the Nansei Islands, Japan. Mycoscience 60: 228-231.

Poli A, Bovio E, Verkley G, Prigione V, Varese GC (2019) Elbamycella rosea gen. et sp. nov. (Juncigenaceae, Torpedosporales) isolated from the Mediterranean Sea. MycoKeys 55: 15-28.

Soowannayana C, Tejab DNC, Yatipa P, Mazumderb FY, Krataitong K, Unagula P, Suetronga S, Preedanona S, Klaysubana A, Sakayaroja J, Sangtieane T (2019) Vibrio biofilm inhibitors screened from marine fungi protect shrimp against acute hepatopancreatic necrosis disease (AHPND). Aquaculture 499: 1–8

Xua W, Gaoa YH, Gonga KF, Lid M, Pang KL (2019) Fungal diversity in the deep-sea hadal sediments of the Yap Trench by cultivation and high throughput sequencing methods based on ITS rRNA gene. Deep-Sea Research Party 1 145: 125-136.



Abdel-Wahab MA, El-Samawaty AEMA, El-Gorbani AM, Yassin AM, Alsaadi MH (2018). Khaleijomyces marinus gen. et sp. nov. (Juncigenaceae, Torpedosporales) a new lignicolous marine fungus from Saudi Arabia. Phytotaxa 340 (3): 277–285.

Devadatha B, Sarma VV (2018). Pontoporeia mangrovei sp. nov, a new marine fungus from an Indian mangrove along with a new geographical and host record of Falciformispora lignatilis. Current Research in Environmental & Applied Mycology. 8(2): 238–246. Doi 10.5943/cream/8/2/8

Devadatha B, Sarma VV, Ariyawansa HA, Jones EBG (2018) Deniquelata vittalii sp. nov., a novel Indian saprobic marine fungus on Suaeda monoica and two new records of marine fungi from Muthupet mangroves, East coast of India. Mycosphere 9(3): 565–582,Doi 10.5943/mycosphere/9/3/8

Devadatha B, Sarma VV, Jeewon R, Wanasinghe DN, Hyde KD, Jones EBG (2018). Thyridariella, a novel marine fungal genus from India: morphological characterization and phylogeny inferred from multigene DNA sequence analyses. Mycological Progress (in press)

Xua W, Lin-feng Gonga LF, Pang KL, Zhu-Hua Luoa ZH (2018) Fungal diversity in deep-sea sediments of a hydrothermal vent system in the Southwest Indian Ridge. Deep-Sea Research Part I 131: 16–26.



Abdel-Wahab MA, Dayarathne MC, Suetrong S, Guo SY, Alias SA, Bahkali AH, Nagahama T, Elgorban AM, Abdel-Aziz FA, Hodhod MS, Al-Hebshi MO, Hyde KD, Nor NABM, Pang* KL and Jones EBG (2017) New saprobic marine fungi and a new combination. Botanica Marina DOI 10.1515/bot-2016-0118.

Azevedo E, Barata M, Marques MI, Caeiro MF (2017) Lulworthia atlantica: a new species supported by molecular phylogeny and morphological analysis. Mycologia 109:2, 287-295, DOI: 10.1080/00275514.2017.1302255

Gnavi G, Garzoli L, Poli A, Prigione V, Burgaud G, Varese GC (2017) The culturable mycobiota of Flabellia petiolata: First survey of marine fungi associated to a Mediterranean green alga. PLoS ONE 12(4): e0175941.

Jones EBG, Ju WT, Lu CL, Guo SY, Ka-Lai Pang KL (2017) The Halosphaeriaceae revisited. Botanica Marina DOI 10.1515/bot-2016-0113.

R. Sridhar (2017). Marine Filamentous Fungi: Diversity, Distribution and Bioprospecting. In: T. Satyanarayana et al. (eds.), Developments in Fungal Biology and Applied Mycology, pp. 59-73.

Krichuk NN, Pivikin MV, Matveeva TV (2017). Three new Penicillium species from marine subaqueous soils. Mycological Progress 16 (1): 15-26.

Nasr Ah, Mohammadimehr M, Vaghei MG, Amoozegar MA, Fazeli SAS, Yurkov A (2017). Jaminaea pallidilutea sp. nov. (Microstromatales), a basidiomycetous yeast isolated from plant material of mangrove forests in Iran. Int J Syst Evol Microbiol 67: 4405–4408. DOI 10.1099/ijsem.0.002302 

Preedanon S, Klaysban A, Suetrong S, Promchoo W, Gundool W, Sangtiean T, Sakayaroj J (2017) Helicascus mangrovei sp.nov. a new intertidal mangrove fungus from Thailand. Mycoscience 58: 174-180.

Raghukumar, S. (2017) Fungi in coastal and oceanic marine ecosystems. Springer International Publishing AG. 378 pp.

Xu W, Guoa S, Pang KL, Luoa ZH (2017) Fungi associated with chimney and sulfide samples from a South Mid-Atlantic Ridge hydrothermal site: Distribution, diversity and abundance. Deep–Sea Research Part I 123: 48–55



Francis MM, Webb V, Zuccarello GC (2016) Marine yeast biodiversity on seaweeds in New Zealand waters. New Zealand J. Bot. 54(1): 30-47.

Xu W, Luo ZH, Guo S, Pang KL (2016) Fungal community analysis in the deep-sea sediments of the Pacific Ocean assessed by comparison of ITS,18Sand28S ribosomal DNA regions. Deep-Sea Research I 10: 951–60.



Xu W, Pang KL, Luo ZH (2014) High Fungal Diversity and Abundance Recovered in the Deep-Sea Sediments of the Pacific Ocean. Microb Ecol 68:688–698 DOI 10.1007/s00248-014-0448-8.


Special issue: Recent advances in marine mycology / Issue editors: Ka-Lai Pang and E.B. Gareth Jones

Unagul, Panida / Suetrong, Satinee / Preedanon, Sita / Klaysuban, Anupong / Gundool, Wunna / Suriyachadkun, Chanwit / Sakayaroj, Jariya (2017) Isolation, fatty acid profiles and cryopreservation of marine thraustochytrids from mangrove habitats in Thailand. Botanica Marina 60: 363-380.

Velmurugan, Natarajan / Lee, Hyun-Mi / Cha, Hyo-Jung / Lee, Yang-Soo (2017) Proteomic analysis of the marine-derived fungus Paecilomyces sp. strain SF-8 in response to polycyclic aromatic hydrocarbons. Botanica Marina 60: 381-392.

Zhong, Tian-Hua / Zhang, Jia-Wei / Xu, Wei / Huang, Liying / Luo, Zhu-Hua (2017) Evaluating the effect of medium composition and fermentation conditions on microbial oil production by a Fusarium strain isolated from the open ocean. Botanica Marina 60: 393-400.

Gleason, Frank H. / Lilje, Osu / Dang, Cecile / Geraci-Yee, Sabrina / Collier, Jackie L. (2017) Possible impacts of zoosporic parasites in diseases of commercially important marine mollusc species: part I. Perkinsozoa (2017). Botanica Marina 60: 401-408.

Collier, Jackie L. / Geraci-Yee, Sabrina / Lilje, Osu / Gleason, Frank H. (2017) Possible impacts of zoosporic parasites in diseases of commercially important marine mollusc species: part II. Labyrinthulomycota. Botanica Marina 60: 409-418.

Scholz, Bettina / Vyverman, Wim / Küpper, Frithjof C. / Ólafsson, Halldór G. / Karsten, Ulf (2017) Effects of environmental parameters on chytrid infection prevalence of four marine diatoms: a laboratory case study. Botanica Marina 60: 419-432.

Rämä, Teppo / Hassett, Brandon T. / Bubnova, Ekaterina (2017) Arctic marine fungi: from filaments and flagella to operational taxonomic units and beyond. Botanica Marina 60: 433-452.

Supaphon, Preuttiporn / Phongpaichit, Souwalak / Sakayaroj, Jariya / Rukachaisirikul, Vatcharin / Kobmoo, Noppol / Spatafora, Joseph W. (2017) Phylogenetic community structure of fungal endophytes in seagrass species. Botanica Marina 60: 489-502. 

Suetrong, Satinee / Preedanon, Sita / Klaysuban, Anupong / Gundool, Wunna / Unagul, Panida / Sakayaroj, Jariya / Promchu, Waratthaya / Sangtiean, Tanuwong (2017) Distribution and occurrence of manglicolous marine fungi from eastern and southern Thailand. Botanica Marina 60: 503-514.


Recent higher order placements of marine fungi

Three recent papers on the higher order ranking of fungi are of interest to marine mycology, although the recommendations are not that clear!

Hyde KD, S. N. Maharachchikumbura, Sinang Hongsanan, Samarakoon MC, Lucking R, Pem D., Harishchandra D, Jeewon R, Zhao RL, Xu JC, Kiu JK, Al-Sadi AM, Bahkali AH, Elgorban AM (2017) The ranking of fungi: a tribute to David L Hawksworth on this 70th birthday. Fungal Diversity 84(3): 1-24.

Sinang Hongsanan, Sajeewa S. N. Maharachchikumbura, Kevin D. Hyde et al. (2017) An updated phylogeny of Sordariomycetes based on phylogenetic and molecular clock evidence. Fungal Diversity 84 (3): 25-41.  

Jian-Kui Liu, Kevin D. Hyde, Rajesh Jeewon, Alan J. L. Phillips et al. (2017) Ranking higher taxa using divergence times: a case study in Dothideomycetes. Fungal Diversity 84 (3): 75-99.


Accepted taxa:


Tirisporellaceae, Tirisporallales, divergence time 112 MYA and share a common ancestor with Sydowiellaceae and Pseudevalsariaceae, but in the phylogenetic tree it is more closely related to Pseudevalsariaceae. The placement of Tirisporellales is not well supported in both MCC and phylogenetic trees and falls within the Diaporthales. However, because it is a marine family and has a hyphomycetous asexual morph. (which is not found in the Diaporthales) it is not synonymised with Diaporthales, until more data are available.

Halosphaeriaceae, Microascales. No changes suggested.

Torpedosporales no changes suggested in Hongsana et al.

Oxydothidaceae, Xylariales, Xylariomycetidae new family for Oxydothis species, many of which are marine and on palm leaves.

Savoryellaceae, Savoryellales New sub class: Savoryeliomycetidae is introduced with three orders: Conioscyphales, Pleurtheciales and Savoryllales.

Lulworthiaceae, Lulworthiales. No changes suggested.



All the following families/orders seem to be supported


Halojulellaceae (weak support)




Salsuginaceae (weak support)


Caryosporaceae (to include Acrocordiopsis)






Wiesneriomycetaceae, Wieseriomycetales

In Hyde et al. (2017), the new order Etheirophorales was proposed for the families Etheirophoraceae and Juncigenaceae, but that should be made if only if the evolution tree is followed. The data is rationalised in the Hongsanan et al. (2017) paper to included phylogenetic and morphological evidence and thus only the changes in Hongsanan et al. should be followed, not the ones in Hyde et al.!


Mushroom Research Foundation 
Chiang Rai 


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