Recent Species

Genus: Ajigaurospora
Class: Sordariomycetes Subclass: Hypocreomycetidae
Order: Microascales
Family: Halosphaeriaceae
pseudopulchella

Genus: Lanspora
Class: Sordariomycetes
Order: Phomatosporales
Family: Phomatosporaceae
dorisauae

Genus: Calophoma
Class: Dothideomycetes
Order: Pleosporales
Family: Didymellaceae
sandfjordenica

Genus: Paralulworthia
Class: Sordariomycetes Subclass: Lulworthiomycetidae
Order: Lulworthiales
Family: Lulworthiaceae
posidoniae

Genus: Paralulworthia
Class: Sordariomycetes Subclass: Lulworthiomycetidae
Order: Lulworthiales
Family: Lulworthiaceae
gigaspora

Genus: Annabella
Class: Leotiomycetes
Order: Helotiales
Family: Cordieritidaceae
australiensis

Genus: Roussoella
Class: Dothideomycetes
Order: Pleosporales
Family: Thyridariaceae, Subclass Pleosporomycetidae
nitidula

Genus: Pontoporeia
Class: Dothideomycetes
Order: Pleosporales
Family: Halotthiaceae, Subclass Pleosporomycetidae
biturbinata

Genus: Neocamarosporium
Class: Dothideomycetes
Order: Pleosporales
Family: Neocamarosporiaceae Wanas., Wijayaw., Crous & K.D. Hyde, Studies in Mycology 87: 245 (2017), Subclass Pleosporomycetidae
phragmitis

Genus: Morosphaeria
Class: Dothideomycetes
Order: Pleosporales
Family: Morosphaeriaceae, Subclass Pleosporomycetidae
muthupetensis

Higher Classification - Section 9

Section 9
MARINE YEASTS: BASIDIOMYCOTA and ASCOMYCOTA

Updated 10 June 2025

Phylum: ASCOMYCOTA

The ascomycetous yeasts are currently undergoing revision and we follow MycoBank in this website.

Subphylum: Saccharomycotina
Class: Saccharomycetes (G. Winter, Rabenh. Krypt.-Fl., Edn 2 (Leipzig) 1.1: 32 (1880)

1. SACCHAROMYCETALES Kudryavtsev, System. Hefen (Berlin): 270 (1960)

Debaryomycetaceae Kurtzman & M. Suzuki, Mycoscience 51(1): 12 (2010)

Candida Berkhout, De schimmelgeslachten Monilia, Oidium, OosporaenTorula: 41 (1923)

This genus is highly polyphyletic and taxonomic changes can be expected.

  1. #C. aaseri Dietrichson ex Uden & H.R. Buckley, in Lodder, Yeasts, a taxonomic study, 2nd Edn (Amsterdam): 912 (1970) (isolated from mangrove soil, water), reported from Doha Bay, Qatar. Molecular data: ITS, LSU
  2. #C. albicans (C.P. Robin) Berkhout, De Schimmelgesl. Monilia, Oidium, OosporaenTorula, Disset. Ultrecht: 44 (1923) (isolate from marine animals), reported from Doha Bay, Qatar. Molecular data: ITS, LSU
  3. C. akabanensis Nakase, M. Suzuki, M. Takash., Rosadi, Hermosillo & Komag., Microbiol. Culture Coll. 10: 35-43 (1994); seawater and sand
  4. C. amphicis S.O. Suh, N.H. Nguyen & M. Blackw., Mycological Research 109 (9): 1049 (2005) (isolated from mangrove decaying matter, and mangrove water)
  5. #C. andamanensis Am-In, Limtong, Yongman. & Jindam., Int. J. Syst. Evol. Microbiol. 61(2): 459 (2011) isolated from mangrove water
  6. #C. asiatica Limtong, Kaewwich., Am-In, Nakase, C.F. Lee, Yongman. &Srisuk, Antonie van Leeuwenhoek 98 (4): 475-481 (2010) (isolated from mangrove water)
  7. #C. atlantica (Siepmann) S.A. Mey. & Simione, in Meyer & Yarrow, Mycotaxon 66: 100 (1998) isolated from sea water and hydrothermal vents deep sea
  8. #C. berthetii Boidin, Pignal, Mermiér & Arpin, Cahiers de La Maboké 1: 100 (1963) isolated from mangrove water
  9. #C. boidinii C. Ramírez, Microbiol. esp. 6(3): 251 (1953) isolated from mangrove water, reported from from Doha Bay, Qatar. Molecular data: ITS, LSU
  10. C. blattae N.H. Nguyen, S.O. Suh & M. Blackw., Mycologia 99(6): 853 (2008) [2007] sand
  11. C. californica (Mrak & McClung ex K.W. Anderson & C.E. Skinner) F.Y. Bai, Z.W. Wu & V. Robert in Wu, Robert & Bai, FEMS Yeast Res. 6(2): 310 (2006) isolated from mangrove water
  12. #C. conglobata (Redaelli) Cif., in Lodder, Manuale di MicologiaMedica, Edn 2 2: 245 (1960) isolated from mangrove water
  13. #C. diddensiae. (Phaff, Mrak & O.B. Williams) Fell & S.A. Mey. [as 'diddensii'], Mycopath. Mycol. appl. 32: 189 (1967) isolated from marine animals
  14. C. duobushaemuli Cend.-Bueno, Kolecka, Alastr.-Izq., Gómez-López, Cuenc.-Estr. & Boekhout [as 'duobushaemulonii'], in Cendejas-Bueno, Kolecka, Alastruey-Izquierdo, Theelen, Groenewald, Kostrzewa, Cuenca-Estrella, Gómez-López & Boekhout, J. Clin. Microbiol. 50(11): 3646 (2012) seawater
  15. #C. ecuadorensis S.A. James, International Journal of Systematic and Evolutionary Microbiology 63 (1): 393 (2012) (isolated from mangrove decaying organic matter)
  16. #C. freyschussii H.R. Buckley & Uden, Mycopath. Mycol. appl. 36: 263 (1968) isolated from animal subatrates
  17. #C. germanica Kurtzman, Robnett & Yarrow, Antonie van Leeuwenhoek 80(1): 79 (2001) isolated from mangrove water
  18. #C. glabrata (H.W. Anderson) S.A. Mey. & Yarrow, in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 612 (1978) isolated from mangrove water, dead leaves,
  19. #C. glaebosa Komag. & Nakase, J. gen. appl. Microbiol., Tokyo 11: 262 (1965) isolated from marine sediments, and from Antarctic marine invertebrates
  20. #C. guilliermondii (Castell.) Langeron & Guerra, Annls Parasit. hum. comp. 16(5): 467 (1938) isolated from seawater including mangroves
  21. C. haemuli (Uden & Kolip.) S.A. Mey. & Yarrow [as 'haemulonii'], in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 612 (1978) seawater and sand
  22. #C. haemulonis (Uden & Kolip.) S.A. Mey. & Yarrow [as 'haemulonii'], in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 612 (1978)
  23. #C. inconspicua (Lodder & Kreger-van Rij) S.A. Mey. & Yarrow, in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 612 (1978)
  24. C. insectorum D.B. Scott, Van der Walt & Klift, in van der Walt, Scott & van der Klift, Mycopath. Mycol. appl. 47(3): 226 (1972) (isolated from mangrove soil, water decaying matter)
  25. #C. intermedia (Cif. & Ashford) Langeron & Guerra, Annls Parasit. hum. comp. 16(5): 461 (1938) (isolated from mangrove water, decaying matter)
  26. #C. laemsonensis Am-In, Limtong, Yongman. & Jindam., Int. J. Syst. Evol. Microbiol. 61(2): 458 (2011) isolated from mangrove water
  27. #C. maltosa Komag., Nakase & Katsuya, J. gen. appl. Microbiol., Tokyo 10: 327 (1964) (isolated from mangrove soil and sediments)
  28. #C. maris (Uden & Zobell) S.A. Mey. & Yarrow, in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 613 (1978) isolated from seawater
  29. #C. maritima (Siepmann) Uden & H.R. Buckley, in Lodder, Mycotaxon 17: 298 (1983) isolated from mangrove water
  30. #C. melibiosica H.R. Buckley & Uden, Mycopath. Mycol. appl. 36: 264 (1968) isolated from animal substrates
  31. #C. membranifaciens (Lodder & Kreger-van Rij) Wick. & Burton, J. Bact. 68: 597 (1954) isolated from mangrove water
  32. 36. C. mesenterica (A. Geiger) Diddens & Lodder, Die Hefasammlung des 'Centraalbureauvoor Schimmelcultures': Beitragezueiner Monographie der Hefearten. II. Teil. Die anaskosporogenen Hefen. ZweiteHälfte: 196 (1942)
  33. #C. metapsilosis Tavanti, A. Davidson, Gow, M. Maiden & Odds, Journal of Clinical Microbiology 43 (1): 290 (2005) (isolated from deep sea sediments)
  34. #C. michaelii S.O. Suh, N.H. Nguyen & M. Blackw., Mycol. Res. 109(9): 1049 (2005) isolated from mangrove water
  35. C. mogii Vidal-Leir., Antonie van Leeuwenhoek 33: 342 (1967)
  36. #C. natalensis Van der Walt & Tscheuschner, Antonie van Leeuwenhoek 23: 189 (1957) isolated from mangrove water
  37. #C. neustonensis C.F. Chang & S.M. Liu, Antonie van Leeuwenhoek 97(1): 38 (2010)
  38. #C. nonsorbophila Nakase, Jindam., Am-in, Ninomiya, H. Kawas. & Limtong, FEMS Yeast Research 9 (4): 663-667 (2009) (isolated from mangrove water)
  39. C. norvegica (Reiersöl) S.A. Mey. & Yarrow, in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 613 (1978)
  40. #C. oceani Burgaud & Barbier, Antonie van Leeuwenhoek 100 (1): 79 (2011) (isolated from deep hydrothermal vent)
  41. C. oleophila Montrocher, Revue Mycol., Paris 32: 73 (1967)
  42. #C. orthopsilosis Tavanti, A. Davidson, Gow, M. Maiden & Odds, Journal of Clinical Microbiology 43 (1): 290 (2005) (isolated from mangrove water)
  43. #C. palmioleophila Nakase & Itoh, Journal of General and Applied Microbiology Tokyo 34 (6): 496 (1988) Isolated from dead leaves in mangroves isolated from mangrove water; on juvenile turtle
  44. #C. parapsilosis (Ashford) Langeron & Talice, Annls Parasit. hum. comp. 10: 1 (1932) (isolated from deep sea sediments and mangrove water, dead leaves, wide range of substrates, isolated from corals and zoanthids on various taxa, common )
  45. #C. phangngaensis Limtong, Yongman., H. Kawas. & Tats. Seki [as 'phangngensis'], in Limtong, Youngmanitchai, Kawasaki & Seki, Int. J. Syst. Evol. Microbiol. 58(2): 517 (2008) isolated from mangrove water
  46. #C. picinguabensis Ruivo, Pagnocca, Lachance & C.A. Rosa, in Ruivo, Lachance, Rosa, Bacci & Pagnocca, Int. J. Syst. Evol. Microbiol. 56(5): 1149 (2006) isolated from mangrove water
  47. C. pinus (Lodder & Kreger-van Rij) S.A. Mey. & Yarrow, in Yarrow & Meyer, Int. J. Syst. Bacteriol. 28(4): 613 (1978)
  48. #C. pseudointermedia Nakase, Komag. & Fukaz., Journal of General and Applied Microbiology Tokyo 22 (4): 178 (1976) Dead mangrove leaves, isolated from mangrove water
  49. #C. pseudolambica M.T. Sm. & Poot, in Smith, Poot & Kull, Stud. Mycol. 31: 175 (1989) (isolated from mangrove soil, water)
  50. #C. ranongensis Am-In, Limtong, Yongman. & Jindam., Int. J. Syst. Evol. Microbiol. 61(2): 459 (2011) isolated from mangrove water
  51. #C. rhizophorensis Fell, M.H. Gut., Statzell & Scorzetti [as 'rhizophoriensis'], in Fell, Statzell-Tallman, Scorzetti & Gutiérrez, Antonie van Leeuwenhoek 99(3): 545 (2011) isolated from mangrove water
  52. C. rugopelliculosa Nakase (1971) J. gen. appl. Microbiol., Tokyo 17(5). Marine yeasts Qatar
  53. C. saitoana Nakase & M. Suzuki, J. gen. appl. Microbiol., Tokyo 31: 85 (1985)
  54. #C. sake (Saito & M. Ota) Uden & H.R. Buckley ex S.A. Mey. & Ahearn, in Lodder, Mycotaxon 17: 298 (1983) isolated from animal substrates
  55. #C. sanitii Limtong, Am-In, Kaeww., Boonmak, Jindam., Yongman., Srisuk, H. Kawas. & Nakase, in Limtong, Kaewwichian, Am-In, Boonmak, Jindamorakot, Yongmanitchai, Srisuk, Kawasaki & Nakase, FEMS Yeast Res. 10(1): 118 (2010)
  56. C. santamariae Montrocher, Revue Mycol., Paris 32: 77 (1967)
  57. #C. sekii Limtong, Kaeww., Jindam., Am-In, Boonmak, Yongman., Srisuk, H. Kawas. & Nakase, in Limtong, Kaewwichian, Am-In, Boonmak, Jindamorakot, Yongmanitchai, Srisuk, Kawasaki & Nakase, FEMS Yeast Res. 10(1): 121 (2010)
  58. #C. sharkensis Fell, M.H. Gut., Statzell & Scorzetti: 542 (2011) isolated from mangrove water
  59. #C. silvanorum Van der Walt, Klift & D.B. Scott, Antonie van Leeuwenhoek 37 (4): 454 (1971) (isolated from mangrove soil and mangrove water)
  60. #C. sinolaborantium S.O. Suh, N.H. Nguyen & M. Blackw., Mycol. Res. 109 (9): 1053 (2005), reported from Salwa Beach, Qatar. Molecular data: ITS, LSU
  61. C. solani Lodder & Kreger-van Rij, Yeasts, a taxonomic study, [Edn 1] (Amsterdam): 672 (1952)
  62. #C. sorboxylosa Nakase, Journal of General and Applied Microbiology Tokyo 17 (5): 392 (1971) (isolated from mangrove water, decaying organic matter)
  63. #C. spencermartinsiae Statzell-Tallman, Scorzetti & Fell, International Journal of Systematic and Evolutionary Microbiology 60 (8): 1980 (2010) isolated from seawater adjacent to corals, isolated from corals and zoanthids
  64. C. suecica Rodr. Mir. & Norkrans, Antonie van Leeuwenhoek 34: 115 (1968)
  65. #C. suwanaritii Limtong, Boonmak, Kaeww., Am-In, Jindam., Yongman., Srisuk, H. Kawas. & Nakase, in Limtong, Kaewwichian, Am-In, Boonmak, Jindamorakot, Yongmanitchai, Srisuk, Kawasaki & Nakase, FEMS Yeast Res. 10(1): 120 (2010) isolated from mangrove waters
  66. #C. taylorii Statzell-Tallman, Scorzetti & Fell, [as 'taylori'], Int. J. Syst. Evol. Microbiol. 60(8): 1981 (2010) isolated from multiple locations among coral reefs and mangrove habitats
  67. #C. thaimueangensis Limtong, Yongman., H. Kawas. & Tats. Seki, Int. J. Syst. Evol. Microbiol. 57(3): 651 (2007) (isolated from mangrove soil, water, decaying organic matter)
  68. #C. tropicalis (Castell.) Berkhout, De Schimmelgesl. Monilia, Oidium, OosporaenTorula, Disset. Utrecht: 44 (1923) (isolated from mangrove soil, water and decaying matter, isolated from corals and zoanthids, many species)
  69. #C. viswanathii Sandu & H.S. Randhawa, Mycopath. Mycol. appl. 18: 179 (1962) (isolated from mangrove decaying matter)
  70. C. zeylanoides (Castell.) Langeron & Guerra, Annls Parasit. hum. comp. 16(5): 501 (1938) (isolated from corals and zoanthids)

Debaryomyces Lodder & Kreger-van Rij, in Kreger-van Rij, Yeasts, a taxonomic study, 3rd Edn (Amsterdam): 130, 145 (1984)

  1. #D. hansenii (Zopf) Lodder& Kreger, The Yeasts: a taxonomic study: 280 (1952)
  2. D. delbrueckii (Lindner) Kudryavtsev, Sistematika Drozhzhei (Moscow): 309 (1954)
  3. D. nepalensis Goto & Sugiy., J. Jap. Bot. 43: 103 (1968) (isolated from mangrove soil, water)

Kurtzmaniella Lachance & Starmer, Int. J. Syst. Evol. Microbiol. 58(2): 523 (2008)

  1. K. natalensis (Van der Walt & Tscheuschner) C.A. Rosa, Casareg. & Lachance, Int. J. Syst. Evol. Microbiol. 69(5): 1507 (2019)

Lodderomyces Van der Walt, Antonie van Leeuwenhoek 32: 2 (1966)

  1. #L. elongisporus (Recca & Mrak) van der Walt, Bothalia 10(3): 418 (1971) isolated from mangrove water

Meyerozyma Kurtzman & M. Suzuki, Mycoscience 51(1): 8 (2010)

  1. #Me. caribbica (Vaughan-Mart.,Kurtzman, S.A. Mey. & E.B. O'Neill) Kurtzman & M. Suzuki, Mycoscience 51(1): 8 (2010) (isolated from mangrove water, isolated from corals and zoanthids)
  2. #M. carpophila (Phaff & M.W. Mill.) A.M. Yurkov & G. Péter, International Journal of Systematic and Evolutionary Microbiology 67 (10): 3981 (2017) isolated from mangrove waters
  3. #Me. guilliermondii (Wick.) Kurtzman & M. Suzuki, Mycoscience 51(1): 7 (2010) (isolated from deep sea sediments, isolated from mangrove soil, water, decaying matter, isolated from corals and zoanthids)
  4. #Me. neustonensis Yurkov & G. Péter, in Yurkov, Dlauchy & Péter, Int. J. Syst. Evol. Microbiol. 67(10): 3981 (2017) isolated from sand

Millerozyma Kurtzman & M. Suzuki, Mycoscience 51 (1): 8 (2010)

  1. M. farinosa (Lindner) Kurtzman & M. Suzuki, Mycoscience 51 (1): 8 (2010)

Priceomyces M. Suzuki & Kurtzman, in Kurtzman & Suzuki, Mycoscience 51(1): 8 (2010)

  1. #P. carsonii (Phaff & E.P. Knapp) M. Suzuki, in Kurtzman & Suzuki, Mycoscience 51(1): 9 (2010)

Scheffersomyces Kurtzman & M. Suzuki, Mycoscience 51 (1): 10 (2010

  1. S. spartinae (Ahearn, Yarrow & Meyers) Kurtzman & M. Suzuki, Mycoscience 51 (1): 9 (2010)

Schwanniomyces Klöcker, Meddelelserfra Carlsberg Laboratoriet 7: 249 (1909)

  1. Sch. etchellsii (Kreger-van Rij) M. Suzuki & Kurtzman, in Kurtzman & Suzuki, Mycoscience 51(1): 11 (2010)
  2. Sch. polymorphus africanus (Van der Walt, Nakase & M. Suzuki) M. Suzuki & Kurtzman, in Kurtzman & Suzuki, Mycoscience 51(1): 11 (2009)
  3. #Sch. pseudopolymorphus (C. Ramírez & Boidin) M. Suzuki & Kurtzman, Mycoscience 51 (1): 11 (2010) (isolated from mangrove water)
  4. #Sch. vanrijiae (van der Walt & Tscheuschner) M. Suzuki & Kurtzman, Mycoscience 51 (1) (2010) (isolated from mangrove soil, decaying matter, mangrove waters)

Suhomyces M. Blackw. & Kurtzman, FEMS Yeast Research 16 (5): fow 41: 5 (2016)

  1. #S. atakaporum (S.-O. Suh & M. Blackw.) M. Blackw. & Kurtzman, FEMS Yeast Research 16 (5): fow041: 6 (2016) (isolated from mangrove water)
  2. #S. choctaworum (S.O. Suh & M. Blackw.) M. Blackw. & Kurtzman, FEMS Yeast Res. 16(5): 10.1093/femsyr/fow041, 6 (2016), isolated from mangrove water

Yamadazyma Billon-Grand, Mycotaxon 35(2): 202 (1989)

  1. #Y. barbieri Burgaud, M. Coton, N. Jacques, Debaets, Maciel, C.A. Rosa, Gadanho, J.P. Samp. & Casareg., International Journal of Systematic and Evolutionary Microbiology 66: 3605 (2016) (isolated from Mid-Atlantic Ridge hydrothermal seawater)
  2. #Y. mexicana (M. Miranda, Holzschu, Phaff & Starmer) Billon-Grand, Mycotaxon 35(2): 203 (1989) (isolated from mangrove water)

Dipodascaceae Engl. & E. Gilg, Syllabus, Edn 9 & 10 (Berlin): 59 (1924)

Dipodascus Lagerh., Jahrbücher für Wissenschaftliche Botanik 24: 549 (1892)

  1. D. geotrichum (E.E. Butler & L.J. Petersen) Arx, Antonie van Leeuwenhoek 43: 336 (1977)
  2. #D. siamensis (Limtong, Kaewwich., Yongman., Srisuk & Fujiyama) P.M. Kirk, Index Fungorum 270: 1 (2015) (isolated from mangrove water)

Geotrichum Link, Magazin der Gesellschaft Naturforschenden Freunde Berlin 3 (1): 17 (1809)

  1. #G. candidum Link, Magazin der Gesellschaft Naturforschenden Freunde Berlin 3 (1): 17, t. 1:26 (1809), reported from Doha Bay and other sites in Qatar. Molecular data: ITS, LSU
  2. #G. europaeum de Hoog & M.T. Sm., Studies in Mycology 50 (2): 502 (2004), reported from Salwa Beach, Qatar. Molecular data: ITS, LSU
  3. #G. phurueaensis Kaewwichian, Yongmanitchai, Srisuk, Fujiyama & Limtong, FEMS Yeast Research 10 (2): 214-220 (2010) (isolated from mangrove water)

Metschnikowiaceae T. Kamieński ex Doweld, Index Fungorum 33: 1 (2013)

Clavispora Rodr. Mir., Antonie van Leeuwenhoek 45: 480 (1979)

  1. Cl. lusitaniae Rodr. Mir., Antonie van Leeuwenhoek 45(3): 480 (1979)

Kodamaea Y. Yamada, Tom. Suzuki, M. Matsuda &Mikata, Biosc., Biotechn., Biochem. 59(6): 1174 (1995)

  1. #Kod. ohmeri (Etchells & T.A. Bell) Y. Yamada, Tom. Suzuki, M. Matsuda & Mikata, Biosc., Biotechn., Biochem. 59(6): 1174 (1995) (isolated from mangrove waters, isolated from corals and zoanthids)

Kregervanrija Kurtzman, FEMS Yeast Res. 6(2): 289 (2006)

  1. #Kr. fluxuum (Phaff & E.P. Knapp) Kurtzman, FEMS Yeast Res. 6(2): 291 (2006) FEMS Yeast Research 6 (2): 289 (2006)

Metschnikowia Kamienski, Trudy imp. S-peterb. Obshch. Estest.: 364 (1899)

  1. M. australis (Fell & I.L. Hunter) Mend.-Hagler, Hagler, Phaff & Tredick, Canad. J. Microbiol. 31 (10): 908 (1985)
  2. #Met. bicuspidata (Metschn.) T. Kamieński, Trudy S. Petersb. Obschch. Est. Otd. Bot. 30(1): 363 (1900)
  3. #Met. krissii (Uden & Cast.-Branco) Uden, Revta Biol., Lisb. 3: 96 (1962)
  4. #Met. pulcherrima Pitt & M.W. Mill., Mycologia 60(3): 669 (1968)
  5. #Met. reukaufii Pitt & M.W. Mill., Mycologia 60(3): 671 (1968)
  6. M. torresii (Uden & Zobell) Lachance, in Lee, Hsiang & Lachance, Antonie van Leeuwenhoek 111(10): 1952 (2018)
  7. #Met. zobellii (Uden & Cast.-Branco) Uden, Revta Biol., Lisb. 3(1): 96 (1962)

Phaffomycetaceae Y. Yamada, H. Kawas., Nagats., Mikata & Tats. Seki, Bioscience Biotechnology and Biochemistry 63: 831

Barnettozyma Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 948 (2008)

  1. #Bar. californica (Lodder) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 948 (2008)

Cyberlindnera Minter, Mycotaxon 110: 473 (2009)

  1. #Cyb. fabianii (Wick.) Minter, Mycotaxon 110: 474 (2009) isolated from mangrove water
  2. #Cyb. jadinii (Sartory, R. Sartory, Weill & J. Mey.) Minter, Mycotaxon 110: 474 (2009) isolated from mangrove water; marine yeast, Qatar
  3. #Cyb. saturnus (Klöcker) Minter, Mycotaxon 110: 476 (2009) isolated from mangrove water

Starmera Y. Yamada, Higashi, S. Ando & Mikata, Bulletin of the Faculty of Agriculture Shizuoka University 47: 31

  1. S. caribaea (Phaff, Starmer, Lachance, V. Aberdeen & Tredick) Y. Yamada, H. Kawas., Nagats., Mikata & T. Seki, Biosc., Biotechn., Biochem. 63(5): 830 (1999)

Wickerhamomyces Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 951 (2008)

  1. #W. anomalus (E.C. Hansen) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 952 (2008)(isolated from mangrove water, isolated from corals and zoanthids)
  2. #W. bovis (Uden & Carmo Souza) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 952 (2008)
  3. #W. canadensis (Wick.) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 952 (2008)
  4. #W. hampshirensis (Kurtzman) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 952 (2008)
  5. #W. pijperi (Van der Walt & Tscheuschner) Kurtzman, Robnett & Basehoar-Powers, FEMS Yeast Research 8 (6): 952 (2008)
  6. #W. rabaulensis (Soneda & S. Uchida) Kurtzman, Robnett & Basehoar-Powers, FEMS Yeast Research 8 (6): 952 (2008)
  7. #W. sydowiorum (D.B. Scott & Van der Walt) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 952 (2008) isolated from mangrove water

Pichiaceae Zender, Bull. Soc. bot. Genève, 2 sér. 17: 290 (1925)

Dekkera Van der Walt, Antonie van Leeuwenhoek 30: 278 (1964)

  1. D. bruxellensis Van der Walt, Antonie van Leeuwenhoek 30: 278 (1964), marine yeasts isolated from sweater Qatar

Martiniozyma Kurtzman, Antonie van Leeuwenhoek 108 (4): 807 (2015)

  1. #M. asiatica Limtong, Kaewwich., Am-In, Nakase, C.F. Lee, Yongman. & Srisuk ex Kurtzman, Antonie van Leeuwenhoek 108 (4): 807 (2015)

Ogataea Y. Yamada, K. Maeda & Mikata, Bioscience Biotechnology and Biochemistry 58 (7): 1253 (1994)

  1. O. angusta (Teun., H.H. Hall & Wick.) Suh & Zhou, FEMS Yeast Research 10 (5): 635 (2010)

Pichia E.C. Hansen, Zentbl. Bakt. ParasitKde, Abt. II 12(19): 538 (1904)

  1. P. fermentans Lodder, Zentbl. Bakt. ParasitKde, Abt. II 86: 242 (1932)
  2. #P. kluyveri Bedford ex Kudryavtsev, Bot. Mater. Otd. Sporov. Rast. Bot. Inst. KomarovaAkad. Nauk S.S.S.R. 13: 145 (1960
  3. #P. kudriavzevii Boidin, Pignal & Besson, Bull. trimest. Soc. mycol. Fr. 81(4): 589 (1965) (isolated from mangrove water)
  4. P. mandshurica Saito, Report of the Central Laboratory, South Manchuria Railway Company 1: 35 (1914)
  5. P. membranifaciens (E.C. Hansen) E.C. Hansen, Zentbl. Bakt. ParasitKde, Abt. II 12(19-21): 538 (1904)
  6. P. norvegensis Leask & Yarrow, Sabouraudia 14: 61 (1976)
  7. P. occidentalis (Kurtzman, M.J. Smiley & C.J. Johnson) Kurtzman, Robnett & Bas.-Powers, FEMS Yeast Res. 8(6): 946 (2008) (isolated from mangrove water)
  8. #P. sporocuriosa G. Péter, Tornai-Leh., Dlauchy & Vitányi, Antonie van Leeuwenhoek 77: 39 (2000) (isolated from mangrove water)
  9. P. terricola Van der Walt, Antonie van Leeuwenhoek 23: 28 (1957)

Saturnispora Z. Liu & Kurtzman, Antonie van Leeuwenhoek 60: 28 (1991)

  1. #S. mangrovi El-Samawaty, Boekhout & Abdel-Wahab, International Journal of Systematic and Evolutionary Microbiology 10.1099/ijsem.0.003859: 4 (2019)
  2. S. mendoncae Kurtzman, FEMS Yeast Res. 6(2): 292 (2006)
  3. #S. sanitii Limtong, Am-In, Kaewwich., Boonmak, Jindam., Yongman., Srisuk, H. Kawas. & Nakase ex Kurtzman, Antonie van Leeuwenhoek 108 (4): 808 (2015)
  4. S. satoi (K. Kodama, Kyono & S. Kodama) Z.W. Liu & Kurtzman, Antonie van Leeuwenhoek 60(1): 28 (1991)
  5. #S. sekii Limtong, Kaewwich., Jindam., Am-In, Boonmak, Yongman., Srisuk, H. Kawas. & Nakase ex Kurtzman, Antonie van Leeuwenhoek 108 (4): 808 (2015) isolated from mangrove water
  6. #S. silvae (Vidal-Leir. & Uden) Kurtzman, Antonie van Leeuwenhoek 108(4): 808 (2015)
  7. #S. suwanaritii Limtong, Boonmak, Kaewwich., Am-In, Jindam., Yongman., Srisuk, H. Kawas. & Nakase ex Kurtzman, Antonie van Leeuwenhoek 108 (4): 808 (2015

Saccharomycetaceae G. Winter, Rabenh. Krypt.-Fl., Edn 2 (Leipzig) 1.1: 58 (1880)

Citeromyces Santa María, Bol. Inst. Nac. Invest. Agron. 17: 275 (1957)

  1. Cit. matritensis (Santa María) Santa María, Bol. Inst. Invest. Agron. Madr. 17(no. 37): 275 (1957)

Grigorovia Gouliamova & Dimitrov, Comptes Rendus de l'Académie Bulgare des Sciences 73(1): 55 (2020)

  1. G. yakushimaensis (Mikata & Ueda-Nishim.) Gouliamova & Dimitrov, Comptes Rendus de l'Académie Bulgare des Sciences 73(1): 55 (2020)

Kazachstania Zubcova, Bot. Mater. Gerb. Inst. Bot. Akad. Naukkazakh. SSR 7: 53 (1971)

  1. #K. bovina Kurtzman & Robnett, J. Clin. Microbiol. 43(1): 105 (2005)
  2. #K. exigua (Reess ex E.C. Hansen) Kurtzman, FEMS Yeast Res. 4(3): 238 (2003)
  3. #K. heterogenica Kurtzman & Robnett, J. Clin. Microbiol. 43(1): 107 (2005)
  4. #K. humilis (E.E. Nel & Van der Walt) N. Jacques, Sarilar & Casarég in Jacques, Sarilar, Urien, Lopes, Morais, Uetanabaro, Tinsley, Rosa, Sicard & Casaregola, Int. J. Syst. Evol. Microbiol. 66(12): 5199 (2016)
  5. #K. jiainica C.F. Lee & Chun H. Liu, FEMS Yeast Res. 8(1): 116 (2008)
  6. #K. pintolopesii Kurtzman, Robnett, J.M. Ward & T.J. Walsh, J. Clin. Microbiol. 43(1): 108 (2005)
  7. K. servazzii (Capr.) Kurtzman (2003) FEMS Yeast Res. 4(3): 238
  8. #K. siamensis Limtong, Yongman., Tun, H. Kawas. & Tats. Seki, Int. J. Syst. Evol. Microbiol. 57(2): 421 (2007)
  9. #K. slooffiae Kurtzman & Robnett, J. Clin. Microbiol. 43(1): 109 (2005)

Kluyveromyces van der Walt, Antonie van Leeuwenhoek 22: 271 (1956)

  1. #Kl. aestuarii (Fell) van der Walt, Antonie van Leeuwenhoek 31: 347 (1965) (isolated from mangrove soil)
  2. Kl. lactis (Stell.-Dekk.) Van der Walt, Bothalia 10(3): 417 (1971)
  3. Kl. lactis var. lactis (Dombr.) van der Walt, Bothalia 10(3): 417 (1971)
  4. Kl. marxianus (E.C. Hansen) van der Walt, Bothalia 10(3): 418 (1971)
  5. Kl. nonfermentans Nagah., Hamam., Nakase &Horikoshi, Int. J. Syst. Bacteriol. 49(4): 1903 (1999)
  6. #Kl. siamensis Am-in, Yongman. & Limtong, FEMS Yeast Research 8 (5): 825 (2008) Am-in, Yongman. & Limtong, FEMS Yeast Research 8 (5): 825 (2008) Isolated from mangrove water, soil, decaying organic matter)

Lachancea Kurtzman, FEMS Yeast Res. 4(3): 239 (2003)

  1. #L. fermentati Kurtzman, FEMS Yeast Res. 4(3): 240 (2003) Isolated from mangrove water
  2. L. kluyveri (Phaff, M.W. Mill. & Shifrine) Kurtzman, FEMS Yeast Res. 4(3): 240 (2003) isolated from sand
  3. #L. meyersii Fell, Statzell & Kurtzman, Stud. Mycol. 50(2): 360 (2004) collected from mangrove habitats in the Bahamas
  4. L. thermotolerans (Filippov) Kurtzman, FEMS Yeast Res. 4(3): 240 (2003)

Saccharomyces Meyen ex Hansen, Vergleichende Morphologie und Biologie der Pilze, Mycetozen und Bacterien: 29 (1883)

  1. S. bayanus Sacc., Sylloge Fungorum 11: 457 (1895)
  2. #S. cerevisiae Meyen ex E.C. Hansen, Meddn Carlsberg Lab. 2: 29 (1883)
  3. S. pastorianus Reess ex E.C. Hansen, Zentbl. Bakt. ParasitKde, Abt. II 12(19-21): 538 (1904)

Saccharomycodaceae Kudryavtsev, System. Hefen (Berlin): 270 (1960)

Hanseniaspora Zikes, Zentbl. Bakt. ParasitKde, Abt. II 30: 148 (1911)

  1. H. guilliermondii Pijper, Verhandelingen Koninklijke Nederlandse Akademie van Wetenschappen Afdeling Natuurkunde 37: 989 (1928) isolated from seawater
  2. H. occidentalis M.T. Sm., Antonie van Leeuwenhoek 40(3): 441 (1974)
  3. H. opuntiae Čadež, Poot, Raspor & M.T. Sm., Int. J. Syst. Evol. Microbiol. 53(5): 1679 (2003) isolated from sand
  4. #H. uvarum (Niehaus) Shehata, Mrak & Phaff ex M.T. Sm., Yeasts, a taxonomic study, 3rd Edn (Amsterdam): 159 (1984), reported from Doha Bay, Qatar. Molecular data: ITS, LSU
  5. H. valbyensis Klöcker, Zentbl. Bakt. ParasitKde, Abt. II 35: 385 (1912)
  6. H. vineae Van der Walt & Tscheuschner, Transactions of the British Mycological Society 40 (2): 212 (1957)

Saccharomycopsidaceae Arx & Van der Walt, Studies in Mycology 30: 174 (1987)

Ambrosiozyma Van der Walt, Mycopathologia et MycologiaApplicata 46 (4): 308 (1972)

  1. A. monospora (Saito) Van der Walt, Mycopathologia et MycologiaApplicata 46 (4): 308 (1972) (isolated from mangrove water)

Saccharomycopsis Schiønning, Meddelelserfra Carlsberg Laboratoriet 6: 93 (1903)

  1. S. crataegensis Kurtzman & Wick., Antonie van Leeuwenhoek 39: 83 (1973)
  2. #S. fibuligera (Lindner) Klöcker, Die Gärungsorganismen in der Theorie und Praxis der Alkoholgarungsgewerbe: 299 (1924), reported from Doha Bay, Qatar. Molecular data: ITS, LSU

Torulaspora Lindner, Jb. Vers.-Lehranst. Brau. Berl. 7: 441 (1904)

  1. T. globosa (Klöcker) van der Walt & Johannsen, C.S.I.R. Res. Rep. 325: 15 (1975)
  2. #T. maleeae Limtong, Imanishi, Jindam., S. Ninomiya, Yongman. & Nakase, FEMS Yeast Res. 8(2): 340 (2008)

Zygotorulaspora Kurtzman, FEMS Yeast Res. 4(3): 243 (2003)

  1. Z. florentina (T. Castelli ex Kudryavtsev) Kurtzman, FEMS Yeast Res. 4(3): 243 (2003)

Trichomonascaceae Kurtzman & Robnett, FEMS Yeast Res. 7(1): 150 (2007)

Blastobotrys Klopotek, Archiv für Mikrobiol 58: 92 (1967)

  1. #Bl. parvus (Fell & Statzell) Kurtzman & Robnett, FEMS Yeast Res. 7(1): 149 (2007)

Groenewaldozyma Kurtzman, Antonie van Leeuwenhoek 109(7): 1042 (2016)

  1. 1. G. salmanticensis (Santa María) Kurtzman, Antonie van Leeuwenhoek 109(7): 1043 (2016)

Starmerella C.A. Rosa & Lachance, Int. J. Syst. Bacteriol. 48(4): 1414 (1998)

  1. S. etchellsii (Lodder & Kreger-van Rij) C.A. Rosa & Lachance, Int. J. Syst. Evol. Microbiol. 68 (4): 1341 (2018)
  2. S. magnoliae (Lodder & Kreger-van Rij) C.A. Rosa & Lachance, Int. J. Syst. Evol. Microbiol. 68(4): 1341 (2018)
  3. S. stellata (Kroemer & Krumbholz) C.A. Rosa & Lachance, Int. J. Syst. Evol. Microbiol. 68(4): 1342 (2018)

Trichomonascus H.S. Jacks., Mycologia 39(6): 712 (1947)

  1. T. ciferrii (M.T. Sm., Van der Walt & Johannsen) Kurtzman & Robnett, FEMS Yeast Res. 7(1): 149 (2007)

Wickerhamiella Van der Walt & Liebenberg, Antonie van Leeuwenhoek 39 (1): 126 (1973)

  1. W. galacta (Golubev & Babeva) C. Vega & Lachance, FEMS Yeast Research 17 (5): fox054, 8 (2017)
  2. #W. martinezcrusiae N.O.P. Maciel, A.R.O. Santos, C.R. Felix, Landell, D.M. Pagani, Pimenta, P. Morais, Angchuan, Wongpanit, Srisuk, Lachance & C.A. Rosa, International Journal of Systematic and Evolutionary Microbiology 71 (11, no. 5092): 3 (2021); beach sandWic
  3. W. pararugosa (Nakase, Komag. & Fukaz.) de Vega & Lachance, FEMS Yeast Res. 17(5): [1] (2016) isolated from sand
  4. W. versatilis (Etchells & T.A. Bell) de Vega & Lachance, FEMS Yeast Res. 17(5): [1] (2016)

Zygoascus M.T. Sm., Antonie van Leeuwenhoek 52: 27 (1986)

  1. Z. hellenicus M.T. Sm., Antonie van Leeuwenhoek 52(1): 27 (1986)

Trigonopsidaceae Lachance & Kurtzman, Int. J. Syst. Evol. Microbiol. 63(8): 3113 (2013)

Trigonopsis Schachner, Zeitschriftfür das GesammteBrauwesen 52: 137 (1929)

  1. Tri. cantarellii (Van der Walt & Kerken) Kurtzman & Robnett, FEMS Yeast Res. 7(1): 150 (2007)

SACCHAROMYCETALES genera incertae sedis

Diutina Khunnamw., Jindam., Limtong & Lachance, International Journal of Systematic and Evolutionary Microbiology 65 (12): 4704 (2015

  1. #D. catenulata (Diddens & Lodder) Khunnamw., Jindam., Limtong & Lachance, International Journal of Systematic and Evolutionary Microbiology 65 (12): 4706 (2015), isolated from sand and sea water in beaches of Olinda, Pernambuco state, Brazil
  2. D. neorugosa (Paredes, Deanna A. Sutton, Cano & Guarro) Khunnamw., Lertwatt., Jindam., Limtong & Lachance, Int. J. Syst. Evol. Microbiol. 65(12): 4708 (2015) isolated from sand
  3. D. rugosa (H.W. Anderson) Khunnamw., Jindam., Limtong & Lachance, Int. J. Syst. Evol. Microbiol. 65(12): 4706 (2015)

Hyphopichia Arx & Van der Walt, Antonie van Leeuwenhoek 42(3): 310 (1976)

  1. Hy. burtonii (Boidin, Pignal, Lehodey, Vey & Abadie) Arx & Van der Walt, Antonie van Leeuwenhoek 42(3): 310 (1976)
  2. #Hy. fennica (Sonck & Yarrow) L.R. Ribeiro, M. Groenew., N. Jacques, Casareg., Lachance & C.A. Rosa, Antonie van Leeuwenhoek 110: 992 (2017), isolated from marine invertebrates in a Southeast Brazilian mangrove ecosystem also water and sand
  3. Hy. homilentoma (Van der Walt & Nakase) L.R. Ribeiro, M. Groenew., M.T. Sm., C.A. Lara, Lachance & C.A. Rosa, Antonie van Leeuwenhoek 110 (7): 992 (2017)
  4. Hy. rhagii (Diddens & Lodder) L.R. Ribeiro, M. Groenew., N. Jacques, Casareg., Lachance & C.A. Rosa, Antonie van Leeuwenhoek 110(7): 992 (2017)

Limtongozyma Boontham, Angchuan, Boonmak & Srisuk, Int. J. Syst. Evol. Microbiol. 71(no. 005123): 1 (2021)

  1. L. cylindracea (Koichi Yamada & Machida ex S.A. Mey. & Yarrow) Boontham, Angchuan, Boonmak & Srisuk, Int. J. Syst. Evol. Microbiol. 71(no. 005123): 1 (2021), (isolated from mangrove soil, water)

Metahyphopichia Sipiczki & Pfliegler, International Journal of Systematic and Evolutionary Microbiology 66 (7): 2550 (2016)

  1. #M. laotica Sipiczki, Pfliegler, Safar, P.B. Morais & C.A. Rosa, International Journal of Systematic and Evolutionary Microbiology 66 (7): 2550 (2016)

Middelhovenomyces Kurtzman & Robnett, Antonie van Leeuwenhoek 105(5): 939 (2014)

  1. M. tepae (Grinb.) Kurtzman & Robnett, Antonie van Leeuwenhoek 105(5): 940 (2014)

Nakazawaea Y. Yamada, K. Maeda & Mikata, Biosc., Biotechn., Biochem. 58(7): 1256 (1994)

  1. #N. anatomiae (Zwillenb.) Kurtzman & Robnett, in Kurtzman & Robnett, FEMS Yeast Res. 14: 1034 (2014), isolated from mangrove water
  2. N. holstii (Wick.) Y. Yamada, K. Maeda & Mikata, Biosc., Biotechn., Biochem. 58(7): 1256 (1994)
  3. #N. ishiwadae (Sugiy. & Goto) Kurtzman & Robnett, FEMS Yeast Research 14 (7): 1034 (2014)
  4. N. siamensis Kaewwich. & Limtong, Int. J. Syst. Evol. Microbiol. 64(1): 268 (2014) isolated from sand

Yarrowia Van der Walt & Arx, Antonie van Leeuwenhoek 46: 519 (1980)

  1. Y. divulgata Nagy, Niss, Dlauchy, Arneborg, D.S. Nielsen & G. Péter, Int. J. Syst. Evol. Microbiol. 63(12): 4821 (2013) (nom. inval.); isolated from juvenile turtles
  2. #Y. deformans (Zach) M. Groenew., T.A. Cout. & M.T. Sm., Antonie van Leeuwenhoek 103(5): 1025 (2013); Isolated from juvenile turtles. Molecuar data: ITS, LSU
  3. Y. galli (G. Péter, Dlauchy, Vasdinyei, Tornai-Leh. & T. Deák) Gouliamova, R.A. Dimitrov, M.T. Sm. & M. Groenew., Persoonia 39: 289 (2017)
  4. Y. lipolytica (Wick., Kurtzman & Herman) Van der Walt &Arx, Antonie van Leeuwenhoek 46: 519 (1980)
  5. # Y. hollandica (Knutsen, V. Robert & M.T. Sm.) Gouliam., R.A. Dimitrov, M.T. Sm. & M. Groenew., Persoonia 39: 289 (2017), oil-degrading yeast isolated from the sea surface microlayer

2. SCHIZOSACCHAROMYCETALES O.E. Erikss., Svedskog & Landvik, Systema Ascomycetum 11: 146 (1993)

Schizosaccharomycetaceae Beij. ex Klöcker, Handbuch der Technischen Mykologie, edn 2: 189 (1905)

Schizosaccharomyces Lindner, Wochenschrift für Brauerei 10: 1298 (1893)

  1. Sch. pombe Lindner, Wochenschrift für Brauerei 10: 1298 (1893)

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